1. Biology Department, Trent University, Peterborough, ON K9J 7B8, Canada
2. Ontario Ministry of Northern Development, Mines, Natural Resources and Forestry, Wildlife Research and Monitoring Section, Trent University, Peterborough, ON K9J 7B8, Canada
3. Landscape Science and Technology Division, Environment and Climate Change Canada, Ottawa, ON K1S 5R1, Canada