1. Department of Veterinary Integrative Biosciences, College of Veterinary Medicine and Biomedical Sciences, Texas A&M University , College Station TX 77845, USA
2. Department of Integrative Biology, University of Wisconsin-Madison , Madison, WI 53706, USA
3. Program in Evolutionary Biology, Department of Ecology and Genetics (IEG), Uppsala University , 752 36 Uppsala, Sweden