1. Key Laboratory of Applied Genetics of universities in Fujian Province, Fujian Agriculture and Forestry University , Fuzhou, Fujian, 350002 , P. R. China
2. National Education Ministry Key Laboratory of Plant Genetic Improvement and Comprehensive Utilization, Fujian Agriculture and Forestry University , Fuzhou, Fujian, 350002 , P. R. China
3. Agricultural College, Fujian Agriculture and Forestry University , Fuzhou, Fujian, 350002 , P. R. China
4. College of Horticultural Sciences, Zhejiang Agriculture and Forestry University , Hangzhou, Zhejiang, 350002 , P. R. China
5. Baoji Academy of Agricultural Sciences , Baoji, Shaanxi, 350002 , P. R. China