1. Institute of Crop Sciences, Chinese Academy of Agricultural Sciences (CAAS)/National Key Facility for Crop Gene Resources and Genetic Improvement, Key Laboratory of Biology and Genetic Improvement of Triticeae Crops, Ministry of Agriculture , Beijing 100081 , China
2. College of Agronomy, Sichuan Agricultural University , Chengdu 611130 , China
3. State Key Laboratory of Protein and Plant Gene Research, Peking-Tsinghua Center for Life Sciences, School of Life Sciences, Peking University , Beijing 100871 , China
4. Institute of Hybrid Wheat, Beijing Academy of Agriculture and Forestry Sciences , Beijing 100097 , China