1. Institute of Physical Education , , Yantai 264025, China
2. Ludong University , , Yantai 264025, China
3. Key Laboratory of Bio-Resource and Eco-Environment of Ministry of Education, College of Life Science, Sichuan University , Chengdu 610065, China
4. Department of Agricultural Sciences , , PO Box 27, Helsinki FI-00014, Finland
5. Viikki Plant Science Centre, University of Helsinki , , PO Box 27, Helsinki FI-00014, Finland
6. College of Agriculture and Biotechnology, Zhejiang University , Hangzhou 310058, China