1. Institute of Biology, Department of Marine Biology, Federal Fluminense University, Outeiro São João Batista, s/no., P.O. Box 100.644, Niterói, Rio de Janeiro 24020-150, Brazil
2. Institute of Biosciences, Federal University of the State of Rio de Janeiro, Av. Pasteur, 296, Urca, Rio de Janeiro 22290-240, Brazil