1. Department of Ecology, Evolution, and Behavior, University of Minnesota, St. Paul, MN, United States of America
2. Department of Earth, Environmental, and Planetary Sciences, Case Western Reserve University, Cleveland, OH, United States of America
3. Northern Research Station, USDA Forest Service, Durham, NH, United States of America
4. Department of Sustainable Resources Management, State University of New York College of Environmental Science and Forestry, Syracuse, NY, United States of America
5. Department of Environmental and Forest Biology, State University of New York College of Environmental Science and Forestry, Syracuse, NY, United States of America