1. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
3. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, China
4. Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional/UFRJ, Rio de Janeiro, Brazil
5. College of Earth Sciences, Jilin University, Changchun, China