1. Hunan Agricultural University, College of Horticulture, Changsha, Hunan, China
2. Engineering Research Center for Horticultural Crop Germplasm Creation and New Variety Breeding, Ministry of Education, Changsha, China
3. Hunan Mid-Subtropical Quality Plant Breeding and Utilization Engineering Technology Research Center, Changsha, China
4. Hunan Key Laboratory of Innovation and Comprehensive Utilization, Changsha, China
5. Hunan Agricultural University, College of Oriental Science & Technology, Changsha, China