1. Swiss Federal Institute of Forest, Snow and Landscape Research (WSL), Zürcherstrasse 111, 8903 Birmensdorf, Switzerland;
2. Swiss Federal Institute of Forest, Snow and Landscape Research (WSL), Zürcherstrasse 111, 8903 Birmensdorf, Switzerland
3. Department of Geography, University of Bonn, Meckenheimer Allee 166, 53115 Bonn, Germany