1. Fisheries Division, Japan International Research Center for Agricultural Sciences, Tsukuba, Ibaraki 305-8686, Japan
2. Independent, Jalan Baharu, Perai, Penang 13700, Malaysia
3. Institute for East China Sea Research, Nagasaki University, Nagasaki City, Nagasaki 851-2213, Japan
4. Fisheries Research Institute, Department of Fisheries, Batu Maung, Penang 11960, Malaysia