1. Translational and Clinical Research Institute, Faculty of Medical Sciences, Newcastle University, Newcastle upon Tyne, United Kingdom.
2. Department of Developmental Medicine, King Abdullah International Medical Research Center (KAIMRC), Riyadh, Saudi Arabia.
3. King Saud Bin Abdulaziz University for Health Sciences, Riyadh, Saudi Arabia.
4. NIHR Nottingham Biomedical Research Centre, Nottingham University Hospitals NHS Trust, University of Nottingham, Nottingham, United Kingdom.
5. Nottingham Digestive Diseases Centre, School of Medicine, University of Nottingham, Nottingham, United Kingdom.