1. Departamento de Botânica, Faculdade de Ciências da Universidade do Porto, Rua do Campo Alegre 1181, 4150‐191 Porto, Portugal and CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, Universidade do Porto, Campus Agrário de Vairão, 4485‐661 Vairão, Portugal;
2. Duke University, Department of Biology, Durham, North Carolina 27708 USA
3. Molecular and Environmental Plant Sciences Program, Texas A&M University, College Station, Texas 77840 USA
4. Department of Systematic and Evolutionary Botany, Faculty of Life Sciences, University of Vienna, Rennweg 14, A 1030 Vienna, Austria
5. School of Theoretical & Applied Science, Ramapo College Mahwah, New Jersey 07430 USA