1. U.S. Geological Survey, Pacific Island Ecosystems Research Center, Kīlauea Field Station, P.O. Box 44, Hawai‘i National Park, HI 96718, USA.
2. Hawai‘i Cooperative Studies Unit, University of Hawai‘i at Hilo, P.O. Box 44, Hawai‘i National Park, HI 96718, USA.