1. Marine Science Department, University of Hawai#i at Hilo, 200 West Kawili Street, Hilo, HI 96720-4091, USA.
2. Department of Biology, University of Louisiana at Lafayette, 410 E. St. Mary Boulevard, Billeaud Hall, Lafayette, LA, USA.
3. Tropical Conservation Biology and Environmental Sciences Program, University of Hawai‘i at Hilo, 200 West Kawili Street, Hilo, HI 96720-4091, USA.