1. Department of Biology, University of Hawai‘i at Hilo, Hilo, Hawai‘i, USA.
2. National Park Service, Inventory and Monitoring Program, Pacific Island Network, P.O. Box 52, Hawai‘i Volcanoes National Park, Hawai‘i 96718, USA.
3. National Park Service, P.O. Box 369, Makawao, Haleakalā National Park, Hawai‘i 96768, USA.