1. School of Ocean Sciences, University of Wales, Bangor, Bangor, and Proudman Oceanographic Laboratory, Liverpool, United Kingdom
2. Proudman Oceanographic Laboratory, Liverpool, United Kingdom
3. Centre for Environment Fisheries and Aquaculture Science, Lowestoft, Suffolk, United Kingdom
4. Centro de Investigacion Cientifica y de Estudios Superiores de Ensenada, Tijuana-Ensenada, Mexico