1. School of Aquatic and Fishery Sciences, University of Washington, 1122 NE Boat St, Seattle, WA 98195, USA Email: jmhuie@uw.edu
2. Friday Harbor Laboratories, University of Washington, 620 University Road, Friday Harbor, WA 98250, USA Email: fishguy@uw.edu
3. Department of Biological Sciences, George Washington University, 800 22nd St NW, Suite 6000, Washington, DC 20052, USA