1. South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences/Key Laboratory of Marine Ranching, Ministry of Agriculture and Rural Affairs/Guangdong Provincial Key Laboratory of Fishery Ecology and Environment, Guangzhou, China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China
3. Scientific Observation and Research Field Station of Pearl River Estuary Ecosystem, Guangzhou, China