1. School of Veterinary Medicine and Science, University of Nottingham, Loughborough, Leicestershire, United Kingdom
2. School of Science and Technology, Nottingham Trent University, Nottingham, United Kingdom
3. Moredun Research Institute, Pentlands Science Park, Bush Loan, Penicuik, Midlothian, Scotland
4. Institute of Anatomy, Histology and Embryology, Faculty of Veterinary Medicine, Leipzig University, Leipzig, Germany