1. Fluminense Federal University, Laboratory of Veterinary Bacteriology, Biomedical Institute, Niterói, Rio de Janeiro, Brazil
2. Technological Development Center, Federal University of Pelotas, Capão do Leão, Rio Grande do Sul, Brazil
3. Laboratory of Investigation in Medical Microbiology (LIMM), Paulo de Góes Institute of Microbiology, Federal University of Rio de Janeiro, Rio de Janeiro, Brazil
4. National Reference Center for Leptospirosis, Biology of Spirochetes Unit, Institut Pasteur, Paris, France