1. Department of Veterinary Preventive Medicine, The Ohio State University, Columbus, Ohio, USA
2. Sarah Allison Steffee Center for Zoological Medicine, Cleveland Metroparks Zoo, Cleveland, Ohio, USA
3. Department of Planning, Design, and Natural Resources, Cleveland Metroparks, Fairview Park, Ohio, USA
4. Center for Veterinary Medicine, Office of Research, U.S. Food and Drug Administration, Laurel, Maryland, USA