1. Laboratory for Epithelial Immunobiology, Research Center for Allergy and Immunology, RIKEN, 1-7-22 Suehiro, Tsurumi, Yokohama, Kanagawa 230-0045, Japan
2. Cancer Research Institute, Kanazawa University, 13-1 Takaramachi, Kanazawa, Ishikawa 920-0934, Japan
3. Laboratory of Mammalian Genes and Development
4. Cell Biology and Metabolism Branch, National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland 20892
5. Supramolecular Biology, International Graduate School of Arts and Sciences, Yokohama City University, 1-7-22 Suehiro, Tsurumi, Yokohama, Kanagawa 230-0045, Japan