1. Division of Virology, Department of Microbiology and Immunology, Institute of Medical Science, University of Tokyo, Tokyo, Japan
2. Division of Pulmonary Medicine, Department of Medicine, Keio University School of Medicine, Tokyo, Japan
3. Clinical Research Division, Virus Research Center, National Hospital Organization Sendai Medical Center, Sendai, Japan
4. Department of Special Pathogens, International Research Center for Infectious Diseases, Institute of Medical Science, University of Tokyo, Tokyo, Japan
5. Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin–Madison, Madison, Wisconsin, USA