1. 1National Marine Biodiversity Institute of Korea, Seocheon, Chungnam 33662, KOREA
2. 2National Institute of Ecology, Seocheon, Chungnam 33657, KOREA
3. 3Kangwon National University, Chuncheon, Kangwon 24341, KOREA
4. 4Hanwha Aqua Planet Yeosu, Yeosu 59744, KOREA