1. Papanin Institute of Biology of Inland Waters, Russian Academy of Sciences, Borok, Yaroslavl Province 152742 Russia
2. Federal Public Budgetary Scientific Institution All-Russian Research Institute of Plant Protection, VNIISS, Voronezh Province 396030 Russia
3. Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, Saint Petersburg 199034 Russia
4. Chefchaouen Association of Speleology and Mountain Activities, Chefchaouen, Morocco
5. Chefchaouen Association of Speleology and Mountain Activities, Chefchaouen, Morocco.