1. aLaboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1–1, Nada, Kobe, Hyogo 657–8501, Japan
2. bAgriculture and Agri-Food Canada, Canadian National Collection of Insects, 960 Carling Avenue, Ottawa, Ontario, Canada K1A 0C6
3. cResearch Fellow (DC), Japan Society for the Promotion of Science, Tokyo, Japan Version of Record, published online 12 December 2017; published in print 2 January 2019