1. School of Plant Biology, Faculty of Natural and Agricultural Sciences, University of Western Australia, Crawley, WA 6009, Australia
2. Tasmanian Institute of Agricultural Research, University of Tasmania, New Town Research Laboratories, New Town, TAS 7008, Australia
3. Department of Agriculture and Food, Baron-Hay Court, South Perth, WA 6151, Australia; and School of Plant Biology, Faculty of Natural and Agricultural Sciences, University of Western Australia, Crawley, WA 6009, Australia