1. Department of Biology, College of Science, De La Salle University, Manila, Philippines
2. Plant and Soil Health Research Unit, Center for Natural Resources and Environmental Research, De La Salle University, Manila, Philippines
3. Parma Research and Extension Center, University of Idaho, Parma, ID, U.S.A.
4. Small Grains and Potato Germplasm Research Unit, USDA-ARS, Aberdeen, ID, U.S.A.