1. Dep. of Agricultural and Environmental Biology; Graduate School of Agricultural and Life Science; The Univ. of Tokyo; Bunkyo Tokyo 113-8657 Japan
2. Cornell Univ.; Section of Plant Breeding and Genetics; School of Integrative Plant Sciences; Ithaca NY 14853
3. Division of Basic Research; Institute of Crop Science; National Agriculture and Food Research Organization; Tsukuba Ibaraki 305-8518 Japan
4. JST, PRESTO; Kawaguchi Saitama 332-0012 Japan
5. USDA-ARS; RW Holley Center for Agriculture and Health; Ithaca NY 14853