1. Key Laboratory of Agro‐ecological Processes in Subtropical Region, Institute of Subtropical Agriculture Chinese Academy of Sciences Changsha China
2. Huanjiang Observation and Research Station for Karst Ecosystems Chinese Academy of Sciences Huanjiang China
3. College of Resources and Environment University of Chinese Academy of Sciences Beijing China
4. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, South China Botanical Garden Chinese Academy of Sciences Guangzhou China