1. Guangxi Laboratory on the Study of Coral Reefs in the South China Sea Coral Reef Research Centre of China School of Marine Sciences Guangxi University Nanning China
2. Southern Marine Sciences and Engineering Guangdong Laboratory (Zhuhai) Zhuhai China
3. State Key Laboratory of Loess and Quaternary Geology Institute of Earth Environment Chinese Academy of Sciences Xi'an China
4. South China Sea Institute of Oceanology Chinese Academy of Sciences Guangzhou China
5. State Key Laboratory of Marine Geology Tongji University Shanghai China