1. Biogeosciences, Biological Oceanography; Alfred Wegener Institute for Polar and Marine Research; Am Handelshafen 12,; D-27570; Bremerhaven; Germany
2. Geosciences; University of Bremen; Klagenfurter Str.,; D-28359; Bremen; Germany
3. Leibniz Institute for Marine Sciences; IFM-GEOMAR; Düsternbrooker Weg 20,; D-24105; Kiel; Germany
4. The Natural History Museum; Cromwell Road,; London; SW7 5BD; United Kingdom