1. Institute of Low Temperature Science Hokkaido University Sapporo Japan
2. Atmosphere and Ocean Research Institute The University of Tokyo Tokyo Japan
3. National Institute of Polar Research Tokyo Japan
4. Field Science Center for Northern Biosphere Hokkaido University Sapporo Japan
5. Faculty of Fisheries Sciences Hokkaido University Hakodate Japan
6. Arctic Research Center Hokkaido University Sapporo Japan