1. A. N. Severtsov Institute of Ecology and Evolution RAS, Moscow, 119071, RUSSIA
2. Department of Vertebrate Zoology, Faculty of Biology, Lomonosov Moscow State University, Moscow, 119234, RUSSIA
3. Kamchatka Branch of Pacific Geographical Institute of Far Eastern Branch RAS, Petropavlovsk-Kamchatsky, 683024, RUSSIA
4. Department of Biology, Memorial University of Newfoundland, St. John's, Newfoundland and Labrador, NL A1B 3X9, CANADA