1. University of Jendouba , Laboratory of Silvo-Pastoral Resources , Silvo-Pastoral Institute of Tabarka , BP. 345, Tabarka 8110, Tunisia .
2. University of Monastir , Laboratory of Botany, Cryptogamy and Plant Biology, Department of Pharmaceutical Sciences “A”, Faculty of Pharmacy of Monastir , Avenue Avicenna, 5000- Monastir , Tunisia .
3. University of Carthage , IRESA, Laboratory of Forest Ecology , INRGREF, BP. 10, Ariana 2080, Tunisia .
4. Institute of Bioscience and BioResources (IBBR), Research Council (CNR), Unit of Palermo , Corso Calatafimi 414, 90129 Palermo , Italy .