1. Tokyo Metropolitan Institute of Medical Science, Kamikitazawa, Setagaya-ku, Tokyo, Japan
2. PCND Neuroscience Research Institute, Poway, CA, USA
3. Department of Physiology, Nippon Medical School, Tokyo, Japan
4. Institute of Agrobiological Sciences, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan
5. Division of Neurosciences, National Institute on Aging, National Institutes of Health, Bethesda, MD, USA