1. Major in Advanced Bioscience, Graduate School of Agriculture, Kindai University, 3327-204, Nakamachi, Nara, Nara 631-8505, Japan
2. Department of Bioscience and Biotechnology, Faculty of Bioscience and Biotechnology, Fukui Prefectural University, 4-1-1 Matsuoka-Kenjojima, Eiheiji, Fukui 910-1195, Japan