1. aNatural Resources Institute, University of Greenwich, Central Avenue, Chatham Maritime, Kent, ME4 4TB, UK k.i.tomlins@gre.ac.uk
2. aNatural Resources Institute, University of Greenwich, Central Avenue, Chatham Maritime, Kent, ME4 4TB, UK
3. bEscola Superior de Biotecnologia, Universidade Católica Portuguesa|Porto, Edifício de Biotecnologia, Rua de Diogo Botelho, 1327, 4169-005 Porto, Portugal