1. Department of Marine Sciences, University of Georgia , Athens, GA 30602, USA
2. College of Fisheries and Ocean Sciences, University of Alaska Fairbanks , Fairbanks, AK 99775, USA
3. Complex Carbohydrate Research Center, University of Georgia , Athens, GA 30602, USA
4. Department of Microbiology and Cell Science, University of Florida , Gainesville, FL 32611, USA