1. Laboratory of Molecular and Cellular Pathology, and Laboratory of Comparative Pathology, Graduate School of Hokkaido University N15, W7 Kita‐ku Sapporo 060‐8638 Japan
2. Laboratory of Comparative Pathology, Graduate School of Hokkaido University N15, W7 Kita‐ku Sapporo 060‐8638 Japan
3. CREST, JST Sapporo 060‐8638 Japan
4. Cellular Dynamics Laboratory, RIKEN, Discovery Research Institute Wako Saitama 351‐0198 Japan
5. Department of Pathology, NIHD, Toyama Shinjuku Tokyo 162‐8640 Japan
6. Department of Medical Microbiology, University College Dublin 4 Ireland
7. 21st Century COE Program for Zoonosis Control, Graduate School of Hokkaido University Sapporo 060‐8638 Japan
8. Department of Molecular Biology and Diagnosis, Hokkaido University Research Center for Zoonosis Control Sapporo 060‐8638 Japan