1. State Key Laboratory for Infectious Disease Prevention and Control, National Institute for Viral Disease Control and Prevention, Chinese Centre for Disease Control and Prevention, Beijing 102206, China
2. Collaborative Innovation Centre for Diagnosis and Treatment of Infectious Diseases, Hangzhou 310003, Zhejiang province, China
3. Aix Marseille University, IRD French Institute of Research for Development, EHESP French School of Public Health, EPV UMR_D 190 “Emergence des Pathologies Virales”, Marseille 13385, France
4. Centre for Ecology & Hydrology, Oxford OX1 3SR, United Kingdom