1. GlaxoSmithKline, Clinical Imaging Centre, Hammersmith Hospital, London, UK
2. Division of Neuroscience and Mental Health, Imperial College, London, UK
3. GlaxoSmithKline, Clinical Pharmacology Modelling and Simulation, Verona, Italy
4. GlaxoSmithKline, Neurosciences Centre of Excellence for Drug Discovery, Harlow, UK
5. Department of Engineering Science, University of Oxford, Oxford, UK