1. Laboratory of Chromosome Dynamics, Institute of Molecular and Cellular Biosciences, University of Tokyo, 1‐1‐1 Yayoi Tokyo 113‐0032 Japan
2. Graduate School of Agricultural and Life Science, University of Tokyo, 1‐1‐1 Yayoi Tokyo 113‐0032 Japan
3. Laboratory of Nuclear Signaling, Institute of Molecular and Cellular Biosciences, and University of Tokyo, 1‐1‐1 Yayoi Tokyo 113‐0032 Japan