1. Department of Environmental Biotechnology, Genetic Engineering Institute, Mubarak City for Scientific Research, Burg El Arab, Alexandria, Egypt;
2. Department of Natural Resources Sciences, University of Maryland, College Park, 20742, USA;
3. USDA‐ARS, Beltsville, MD 20705 USA;
4. National Research Center, Agricultural Microbiology Department, Dokki, Cairo, Egypt;
5. Botany Department, Faculty of Science, University of Alexandria, Egypt