1. Department of Wildlife, Fish, and Conservation Biology, University of California, Davis, CA 95616, U.S.A. and Serengeti Wildlife Research Institute, PO Box 661, Arusha, Tanzania, Graduate Group in Ecology, University of California, Davis, CA 95616, U.S.A.
2. , Frankfurt Zoological Society, c/o Tanzania National Parks, PO Box 3134, Arusha, Tanzania
3. , Department of Wildlife, PO Box 63150, Dar es Salaam, Tanzania
4. , Natural Resources Institute, Central Avenue, Chatham Maritime, Kent ME4 4TB, U.K.
5. , Tanzania Wildlife Conservation Monitoring, Conservation Information Centre, PO Box 14935, Arusha, Tanzania
6. 6