1. Department of Zoology, LaTrobe University, Bundoora, Victoria 3083, Australia;
2. Department of Biology, Imperial College, Silwood Park, Ascot SL5 7PY, UK;
3. Department of Agriculture and Fisheries for Scotland, East Craigs, Edinburgh EH12 8NJ, UK; and
4. Entomology and Nematology Department, Rothamsted Experimental Station, Harpenden, Hertfordshire, AL5 2JQ, UK