1. Center for Biomembranes and Lipid Enzymology, Institute of Biomembranes, Department of Biochemistry of Membranes, Utrecht University, Padualaan 8, 3584 CH Utrecht, The Netherlands, Department of Medicinal Chemistry, Utrecht University, Sorbonnelaan 16, 3584 CA Utrecht, The Netherlands, and Department of Chemistry and Biochemistry, University of Arkansas, Fayetteville, Arkansas 72701