1. Graduate School of Environmental and Symbiotic Sciences, Prefectural University of Kumamoto, 3-1-100, Tsukide, Kumamoto 862-8502, Japan, Kitakyushu City Institute of Environmental Sciences, Shin-ike 1-2-1, Tobata, Kitakyushu 804-0082, Japan, and Center for Marine Environmental Studies (CMES), Ehime University, Bunkyo-cho 2-5, Matsuyama 790-8577, Japan