1. Department of Biology, University of Maryland, College Park, Maryland 20740, United States
2. Center for Computational Biology, Flatiron Institute, New York, New York 10010, United States
3. Center for Computational Mathematics, Flatiron Institute, New York, New York 10010, United States
4. Fischell Department of Bioengineering, University of Maryland, College Park, Maryland 20740, United States